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The [[family (biology)|family]] '''Threskiornithidae''' includes 34 [[species]] of large terrestrial and wading [[bird]]s, falling into two subfamilies, the '''[[ibis]]es''' and the '''[[spoonbill]]s'''. It was formerly known as Plataleidae. The spoonbills and ibises were once thought to be related to other groups of long-legged wading birds in the order [[Ciconiiformes]]. A recent study found that they are members of the order [[Pelecaniformes]].<ref>{{cite journal |url=http://www.sciencemag.org/cgi/content/abstract/320/5884/1763 |title=A Phylogenomic Study of Birds Reveals Their Evolutionary History |first=Shannon J. |last=Hackett |coauthors=Rebecca T. Kimball; Sushma Reddy; Rauri C. K. Bowie; Edward L. Braun; Michael J. Braun; Jena L. Chojnowski; W. Andrew Cox; Kin-Lan Han; John Harshman; Christopher J. Huddleston; Ben D. Marks; Kathleen J. Miglia; William S. Moore; Frederick H. Sheldon; David W. Steadman; Christopher C. Witt; Tamaki Yuri; |journal=Science |volume=320 |issue=5884 |pages=1763–1768 |doi=10.1126/science.1157704 |month=June |year=2008 |pmid=18583609}}</ref> In response to these findings, the [[International Ornithological Congress]] (IOC) recently reclassified Threskiornithidae and their sister taxa [[Ardeidae]] under the order [[Pelecaniformes]] instead of the previous order of Ciconiiformes.<ref>"Gill, F. & D. Donsker (Eds). 2010. IOC World Bird Names (version 2.4). Available at http://www.worldbirdnames.org/ [Accessed 29 May 2010].</ref> Whether the two subfamilies are reciprocally [[Monophyly|monophyletic]] is an open question. The South American Checklist Committee's entry for the Threskiornithidae includes the following comment "Two subfamilies are traditionally (e.g., Matheu & del Hoyo 1992) recognized: Threskiornithinae for ibises and Plataleinae for spoonbills; because the main distinction has to do with bill shape, additional information, especially genetic, is required to recognize a major, deep split in the family."<ref>{{cite web |url=http://www.aou.org/checklist/south.php |title=A classification of the bird species of South America |work=South American Classification Committee |publisher=American Ornithologists' Union |accessdate=2009-09-09}} {{Dead link|date=September 2010|bot=H3llBot}}</ref>
The [[family (biology)|family]] '''Threskiornithidae''' includes 34 [[species]] of large terrestrial and wading [[bird]]s, falling into two subfamilies, the '''[[ibis]]es''' and the '''[[spoonbill]]s'''. It was formerly known as Plataleidae. The spoonbills and ibises were once thought to be related to other groups of long-legged wading birds in the order [[Ciconiiformes]]. A recent study found that they are members of the order [[Pelecaniformes]].<ref>{{cite journal |url=http://www.sciencemag.org/cgi/content/abstract/320/5884/1763 |title=A Phylogenomic Study of Birds Reveals Their Evolutionary History |first=Shannon J. |last=Hackett |coauthors=Rebecca T. Kimball; Sushma Reddy; Rauri C. K. Bowie; Edward L. Braun; Michael J. Braun; Jena L. Chojnowski; W. Andrew Cox; Kin-Lan Han; John Harshman; Christopher J. Huddleston; Ben D. Marks; Kathleen J. Miglia; William S. Moore; Frederick H. Sheldon; David W. Steadman; Christopher C. Witt; Tamaki Yuri; |journal=Science |volume=320 |issue=5884 |pages=1763–1768 |doi=10.1126/science.1157704 |month=June |year=2008 |pmid=18583609}}</ref> In response to these findings, the [[International Ornithological Congress]] (IOC) recently reclassified Threskiornithidae and their sister taxa [[Ardeidae]] under the order [[Pelecaniformes]] instead of the previous order of Ciconiiformes.<ref>"Gill, F. & D. Donsker (Eds). 2010. IOC World Bird Names (version 2.4). Available at http://www.worldbirdnames.org/ [Accessed 29 May 2010].</ref> Whether the two subfamilies are reciprocally [[Monophyly|monophyletic]] is an open question. The South American Checklist Committee's entry for the Threskiornithidae includes the following comment "Two subfamilies are traditionally (e.g., Matheu & del Hoyo 1992) recognized: Threskiornithinae for ibises and Plataleinae for spoonbills; because the main distinction has to do with bill shape, additional information, especially genetic, is required to recognize a major, deep split in the family."<ref>{{cite web |url=http://www.aou.org/checklist/south.php |title=A classification of the bird species of South America |work=South American Classification Committee |publisher=American Ornithologists' Union |accessdate=2009-09-09}} {{Dead link|date=September 2010|bot=H3llBot}}</ref>

A study of mitochondrial DNA of the spoonbills plus the [[Sacred Ibis|Sacred]] and [[Scarlet Ibis]]es found that the spoonbills formed a clade with old world genus ''[[Threskiornis]]'', with ''[[Nipponia nippon]]'' and ''[[Eudocimus]]'' as progressively earlier offshoots and more distant relatives, and hence casts doubt on the arrangement of the family into ibis and spoonbill subfamilies.<ref>{{cite journal|author=Chesser, R.Terry; Yeung, Carol K.L.; Yao, Cheng-Te; Tians, Xiu-Hua; Li Shou-Hsien|year=2010|title=Molecular phylogeny of the spoonbills (Aves: Threskiornithidae) based on mitochondrial DNA|journal=Zootaxa|issue= 2603|pages=53-60|issn=1175-5326}}</ref>


==Description and ecology==
==Description and ecology==

Revision as of 12:06, 19 August 2011

Threskiornithidae
Royal Spoonbill
Scientific classification
Kingdom:
Phylum:
Class:
Order:
Family:
Threskiornithidae

Richmond, 1917
Subfamilies

The family Threskiornithidae includes 34 species of large terrestrial and wading birds, falling into two subfamilies, the ibises and the spoonbills. It was formerly known as Plataleidae. The spoonbills and ibises were once thought to be related to other groups of long-legged wading birds in the order Ciconiiformes. A recent study found that they are members of the order Pelecaniformes.[2] In response to these findings, the International Ornithological Congress (IOC) recently reclassified Threskiornithidae and their sister taxa Ardeidae under the order Pelecaniformes instead of the previous order of Ciconiiformes.[3] Whether the two subfamilies are reciprocally monophyletic is an open question. The South American Checklist Committee's entry for the Threskiornithidae includes the following comment "Two subfamilies are traditionally (e.g., Matheu & del Hoyo 1992) recognized: Threskiornithinae for ibises and Plataleinae for spoonbills; because the main distinction has to do with bill shape, additional information, especially genetic, is required to recognize a major, deep split in the family."[4]

A study of mitochondrial DNA of the spoonbills plus the Sacred and Scarlet Ibises found that the spoonbills formed a clade with old world genus Threskiornis, with Nipponia nippon and Eudocimus as progressively earlier offshoots and more distant relatives, and hence casts doubt on the arrangement of the family into ibis and spoonbill subfamilies.[5]

Description and ecology

Members of the family have long, broad wings with 11 primary feathers and about 20 secondaries. They are strong fliers and, rather surprisingly, given their size and weight, very capable soarers. The body tends to be elongated, the neck more so, with rather long legs. The bill is also long, decurved in the case of the ibises, straight and distinctively flattened in the spoonbills. They are large birds, but mid-sized by the standards of their order, ranging from the Dwarf Olive Ibis (Bostrychia bocagei), at 45 cm (18 in) and 450 g (1 lb), to the Giant Ibis (Thaumatibis gigantea), at 100 cm (40 in) and 4.2 kg (9.2 lb).

They are distributed almost worldwide, being found near almost any area of standing or slow-flowing fresh or brackish water. Ibises are also found in drier areas, including landfills. All are diurnal; spending the day feeding on a wide range of invertebrates and small vertebrates: ibises by probing in soft earth or mud, spoonbills by swinging the bill from side to side in shallow water. At night, they roost in trees near water. They are gregarious, feeding, roosting, and flying together, often in formation.

Nesting is colonial in ibises, more often in small groups or singly in spoonbills, nearly always in trees overhanging water, but sometimes on islands or small islands in swamps. Generally, the female builds a large structure out of reeds and sticks brought by the male. Typical clutch size is 2 to 5; hatching is asynchronic. Both sexes incubate in shifts, and after hatching feed the young by partial regurgitation. Two or three weeks after hatching, the young no longer need to be brooded continuously and may leave the nest, often forming creches but returning to be fed by the parents.

Species

Eurasian Spoonbill Platalea leucorodia in India.

FAMILY: THRESKIORNITHIDAE

References

  1. ^ Myers, P. R. "Family Threskiornithidae (ibises and spoonbills)". Animal Diversity Web. University of Michigan. Retrieved 2009-09-09. {{cite web}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
  2. ^ Hackett, Shannon J. (2008). "A Phylogenomic Study of Birds Reveals Their Evolutionary History". Science. 320 (5884): 1763–1768. doi:10.1126/science.1157704. PMID 18583609. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help); Unknown parameter |month= ignored (help)CS1 maint: extra punctuation (link)
  3. ^ "Gill, F. & D. Donsker (Eds). 2010. IOC World Bird Names (version 2.4). Available at http://www.worldbirdnames.org/ [Accessed 29 May 2010].
  4. ^ "A classification of the bird species of South America". South American Classification Committee. American Ornithologists' Union. Retrieved 2009-09-09. [dead link]
  5. ^ Chesser, R.Terry; Yeung, Carol K.L.; Yao, Cheng-Te; Tians, Xiu-Hua; Li Shou-Hsien (2010). "Molecular phylogeny of the spoonbills (Aves: Threskiornithidae) based on mitochondrial DNA". Zootaxa (2603): 53–60. ISSN 1175-5326.{{cite journal}}: CS1 maint: multiple names: authors list (link)