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At the end of the game, Amaterasu encounters Yami, the main antagonist and final boss of the game who resembles a small fish inside a huge sphere, whose design is altered through the different stages of the battle. Yami is also the ruler of the demons. Before battle, he drains Amaterasu of her powers and leaves her as a plain white wolf. Amaterasu regains her powers throughout the fight, but, after the fourth round, Yami destroys them all again and leaves Amaterasu in a near-dead state. However, when Issun gets everyone to believe in Amaterasu before the fifth and final round, she changes into her most powerful form and battles Yami, vanquishing him forever. In the final battle, Yami has a huge clawed hand, which demonstrates the evil which comes from humans' hands. The word "Yami" means "darkness" in Japanese.
Two other characters reappear several times within the quest. [[Minamoto no Yoshitsune|Waka]] appears to Amaterasu several times in the game as a beautiful young [[Fue (flute)|flute]]-playing man in costume resembling a [[tengu]] (dressed like a [[yamabushi]]). He is aware of the goddess's true identity, foretells her future, and at times battles with her. He leads the Tao Troopers whose members Abe and Kamo are based on the two famous [[Onmyōdō|onmyōji]] [[Abe no Seimei]] and [[Kamo no Yasunori]]. Waka's dialogue, dropping [[French language|French]] affectionate terms at times, conveys a sense of familiarity with Amaterasu, as it turns out that Waka is much older than he appears and has walked with Amaterasu on the Celestial Plain hundreds of years ago.<ref>{{cite web | url = https://www.ign.com/articles/2007/02/14/okami-au-review | title = Ōkami AU Review | last = Shea | first = Cam | date = 13 February 2007 | access-date = 15 June 2020 | work = IGN | publisher = IGN Entertainment | archive-date = 1 May 2023 | archive-url = https://web.archive.org/web/20230501022249/https://www.ign.com/articles/2007/02/14/okami-au-review | url-status = live }}</ref> The other is [[Yamata no Orochi|Orochi]], the eight-headed demon and a major villain within the game which the player will encounter several times. Orochi repeatedly has threatened Kamiki village, demanding a sacrifice of a young woman. Each of its eight heads is infused with a different elemental magic power, but the entire demon is susceptible to a special brew of [[sake]] available only at Kamiki Village, allowing Amaterasu to defeat it while in its stupor.<ref name="playtm-review"/> Amaterasu trusts Queen [[Himiko]], the ruler of "Sei-an City", who is killed by one of the demons.
Throughout the game, the player encounters several other characters that are inspired from Japanese folklore.<ref name="Folklore">{{cite book | year=2006 | editor=Capcom Entertainment, Inc. | title=Ōkami instruction manual | pages=34–35 | publisher=Capcom }}</ref>
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