of Cephalonia have all along been extremely active; and no slight amount of toil has been expended in the construction of terraces on the steep sides of the hills. Owing to the thinness of the population, however, but a small proportion of the soil is under cultivation, and the quantity of grain grown in the island is comparatively meagre. The staple is the currant, in the production of which the island surpasses Zante. The fruit is smaller than that of the Morea, and has a peculiar flavour; it finds a market mainly in Holland, Belgium and Germany. The grape vine also is grown, and the manufacture of wine is a rising industry. The olive crop is of considerable importance, and the culture of cotton in the low grounds has been successfully attempted. Manufactures are few and undeveloped, but lace from the aloe fibre, Turkey carpets and basket-work are produced by the villagers, and boats are built at both the principal towns. Of all the seven Ionian islands Cephalonia and Zante are most purely Greek, and the inhabitants display great mental activity.
In the Homeric poems Cephalonia is generally supposed to be mentioned under the name of Same, and its inhabitants, among the subjects of Ulysses, to be designated Cephallenes (see, however, under Ithaca). In the Persian War they took but little part; in the Peloponnesian they sided with the Athenians. The town of Pale was vainly besieged by Philip of Macedon in 218 B.C., because it had supported the Aetolian cause. In 189 B.C. all the cities surrendered to the Romans, but Same afterwards revolted, and was only reduced after a siege of four months. The island was presented by Hadrian to Athens, but it appears again at a later date as “free and autonomous.” After the division of the Roman empire, it continued attached to Byzantium till 1082, when it was captured by Robert Guiscard, who died, however, before he could repress the revolt of 1085. In 1204 it was assigned to Gaius, prince of Tarentum, who accepted the protection of Venice in 1215; and after 1225 it was held along with Santa Maura and Zante by a succession of five counts of the Tocco family at Naples. Formally made over to Venice in 1350 by the prince of Tarentum, it was afterwards captured by the Turks in 1479; but the Hispanico-Venetian fleet under Benedetto Pessaro and Gonsalvo of Cordova effected their expulsion in 1500, and the island continued in Venetian possession till the fall of the republic. For some time it was administered for the French government, but in 1809 it was taken by the British under Cuthbert, Lord Collingwood. Till 1813 it was in the hands of Major de Bosset, a Swiss in the British service, who displayed an industry and energy in the repression of injustice and development of civilization only outdone by the despotic vigour of Sir Charles Napier, who held the same office for the nine years from 1818 to 1827. During the British protectorate the island made undoubted advances in material prosperity, but was several times the scene of political disturbances. It retained longer than the sister islands traces of feudal influence exerted by the landed proprietors, but has been gradually becoming more democratic. Under the Venetians it was divided into eight districts, and an elaborate system of police was in force; since its annexation to Greece it has been broken up into twenty demarchies, each with its separate jurisdiction and revenues, and the police system has been abolished.
Authorities.—A special treatise on the antiquities of Cephalonia was written by Petrus Maurocenus. See Holland’s Travels (1815); Ansted’s Ionian Islands (1863); Viscount Kirkwall’s Four Years in Ionian Islands (1864); Wiebel’s Die Insel Kephalonia; parliamentary papers. Riemann, Recherches archéologiques sur les Iles Ioniennes (Paris, 1879–1880); Partsch, Kephallenia und Ithaka (1890); see also Corfu; Ionian Islands. (E. Gr.)
CEPHALOPODA, the fifth of the classes into which the zoological phylum Mollusca is divided (see Mollusca). The Cephalopoda are mainly characterized by the concrescence of the foot and head. The foot grows forward on each side so as to surround the mouth, the two upgrowths meeting on the dorsal side of the head—whence the name Cephalopoda. The perioral portion of the foot is drawn out into paired arm-like processes; these may be beset with sheathed tentacles or with suckers or hooks, or both. The epipodia are expanded into a pair of muscular lobes right and left, which are bent round towards one another so that their free margins meet and constitute a short tube—the siphon or funnel. The hind-foot is either very small or absent. A distinctive feature of the Cephalopoda is their bilateral symmetry and the absence of anything like the torsion of the visceral mass seen in the Anisopleurous Gastropoda.
The anus, although it may be a little displaced from the median line, is approximately median and posterior. The mantle-skirt is deeply produced posteriorly, forming a large sub-pallial chamber around the anus. By the side of the anus are placed the single or paired apertures of the nephridia, the genital apertures (paired only in Nautilus, in female Octopoda, female Ommatostrephes and male Eledone), and the paired ctenidia. The visceral hump or dome is elevated, and may be very much elongated in a direction almost at right angles to the primary horizontal axis of the foot.
A shell is frequently, but not invariably, secreted on the visceral hump and mantle-skirt. The shell is usually light in substance or lightened by air-chambers in correlation with the free-swimming habits of the Cephalopoda. It may be external or internal, that is, enclosed in folds of the mantle. Very numerous minute pigmented sacs, capable of expansion and contraction, and known as chromatophores, are usually present in the integument. The sexes are separate.
The ctenidia are well developed as paired gill-plumes, serving as the efficient branchial organs (figs. 4, 24).
The vascular system is very highly developed; the heart consists of a pair of auricles and a ventricle (figs. 12, 28). Branchial hearts are formed on the afferent vessels of the branchiae. It is not known to what extent the minute subdivision of the arteries extends, or whether there is a true capillary system.
The pericardium is extended so as to form a very large sac, passing among the viscera dorsalwards and sometimes containing the ovary or testis—the viscero-pericardial sac—which opens to the exterior either directly or through the renal organs. It has no connexion with the vascular system. The renal organs are always paired sacs, the walls of which invest the branchial afferent vessels (figs. 28, 29). They open each by a pore into the viscero-pericardial sac, except in Nautilus. The anal aperture is median and raised on a papilla. Jaws (fig. 6, e) and a radula (fig. 9) are well developed. The jaws have the form of powerful beaks, either horny or calcified (Nautilus), and are capable of inflicting severe wounds.
Cerebral, pleural and pedal ganglia are present, but the connectives are shortened and the ganglia concentrated and fused in the cephalic region. Large special ganglia (optic, stellate and supra-buccal) are developed. Sense-organs are highly developed; the eye exhibits a very special elaboration of structure in the Dibranchiata, and a remarkable archaic form in the nautilus. Otocysts are present in all. The typical osphradium is not present, except in Nautilus, but other organs are present in the cephalic region, to which an olfactory function is ascribed both in Nautilus and in the other Cephalopoda.
Hermaphroditism is unknown in Cephalopoda; male and female individuals always being differentiated. The genital aperture and duct is sometimes single, when it is the left; sometimes the typical pair is developed right and left of the anus. The males of nearly all Cephalopoda have been shown to be characterized by a peculiar modification of the arm-like processes or lobes of the fore-foot, connected with the copulative function. The term hectocotylization is applied to this modification (see figs. 6, 24). Elaborate spermatophores or sperm-ropes are formed by all Cephalopoda, and very usually the female possesses special capsule-forming and nidamental glands for providing envelopes to the eggs (fig. 4, g.n.). The egg is large, and the development is much modified by the presence of an excessive amount of food-material diffused in the protoplasm of the egg-cell. Trochosphere and veliger stages of development are consequently not recognizable.
The Cephalopoda are divisible into two orders, Tetrabranchiata and Dibranchiata, the names of which (due to Sir R. Owen) describe the number of gill-plumes present; but in fact there are several characters, of as great importance as those derived from the gills, by which the members of these two orders are separated from one another.
Order 1. Tetrabranchiata (= Schizosiphona, Tentaculifera).
Characters.—The inrolled lateral margins of the epipodia are not fused, but form a siphon by apposition (fig. 4). The circumoral lobes of the fore-foot carry numerous retractile tentacles, not suckers (fig. 6). There are two pairs of ctenidial gills (hence Tetrabranchiata), and two pairs of renal organs, consequently four renal apertures (fig. 4). The viscero-pericardial chamber opens by two independent apertures to the exterior, and not into the renal sacs. There are two oviducts (right and left) in the female, and two sperm-ducts in the male, the left duct in both sexes being rudimentary. A large external shell, either coiled or straight, is present, and is not enclosed by reflections of the
